This is read by an automated voice. Please report any issues or inconsistencies here. Gramma, a beloved Galápagos tortoise at the San Diego Zoo believed to be 141 years old, was euthanized after ...

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...

The California zoo invited fans of the late Galápagos tortoise to enjoy one of her favorite snacks, a "generous, fruit-filled salad," in her honor San Diego Zoo/I/nstagram Gramma, the Galápagos ...

Gramma at the zoo’s celebration of what’s believed to have been her 138th birthday in 2022. (Image from @SanDiegoZoo on YouTube) Gramma, a Galápagos tortoise that the San Diego Zoo called “a quiet and ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Gramma was born ...

Gramma, a 141-year-old Galapagos tortoise and the San Diego Zoo’s oldest resident, has died. She came from the Bronx Zoo nearly a century ago and was known as the "Queen of the Zoo." Visitors and ...

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died. Gramma was born ...

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galápagos tortoise Gramma, the oldest resident of the San Diego Zoo, has ...

LOS ANGELES — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise Gramma, the oldest resident of the San Diego Zoo, has died.

Gramma, an estimated 141-year-old Galápagos tortoise, died Thursday at the San Diego Zoo, ending a near century-long chapter of what the zoo said was quiet, yet constant ambassadorship to generations ...